| 1 | Harvard University Boston, MA |
| 2 | Massachusetts Institute of Technology Cambridge, MA |
| Stanford University Stanford, CA | |
| 4 | Johns Hopkins University Baltimore, MD |
| University of California--Berkeley Berkeley, CA | |
| 6 | University of California--San Francisco San Francisco, CA |
| 7 | Yale University New Haven, CT |
| 8 | California Institute of Technology Pasadena, CA |
| 9 | Rockefeller University New York, NY |
| 10 | Princeton University Princeton, NJ |
| University of Texas Southwestern Medical Center--Dallas Dallas, TX | |
| 12 | Washington University in St. Louis St. Louis, MO |
